Built in 1915 but hasn't been a real YMCA in years, the YMCA Building reopened in early 2012 (before flooding destroyed the lower level, necessitating more renovations, including adding a new ADA ramp). The building originally had a chapel, swimming pool, and bowling alley before those functions were replaced by other buildings. They also had a barber shop (also replaced by the MSC). Sometime in the mid-2000s, the building was condemned due to structural concerns (been tough to nail down, one source says 2003, one says 2006, and I think I read somewhere about 2005--also, it was offices before 1995) from previous renovations. In 2010, renovations took place to rebuild the rear wing and extensively renovate the front.
There isn't a whole lot of information on its history: I could dig up pictures, but there's no drama like the Memorial Student Center, either (the building is pretty small). As for the digging up pictures part, there's a Flickr page.
The newer photo below was from this page, though Archive.org didn't seem to save the picture.
Updated 2019